I'm currently using the pre-release GNOME 2.14 (well, 2.13.99 or whatever it is at the moment) in Ubuntu Dapper, and just noticed the cutest behaviour. I've never noticed it before, so it may not be in older versions, but on the other hand it's not the sort of thing I'm very likely to do by accident, so it may have been a feature for years.
Open several windows in the same workspace and hover your mouse cursor over the task bar (by default on your bottom panel). Scroll your mouse wheel up and down and observe what happens. Cool, eh? Maybe not useful, but cool. Try the same trick with the workspace switcher.
Anybody know any other non-obvious things you can do with your mouse wheel?
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Changing timezones in KDE
Here's one I just discovered
Open a folder in Nautilus (the GNOME file manager). With the mouse cursor over the icons, hold down 'Control' and scroll the mouse wheel up or down.